TDR Targets

Basic information

Technical information

Name: Unnamed compound
MW: 380.417 | Formula: C20H16N2O4S
H donors: 2 H acceptors: 4 LogP: 4.03 Rotable bonds: 5
Rule of 5 violations (Lipinski): 1
SMILES: C=CCN1/C(=N/c2cccc(c2)C(=O)O)/S/C(=C\c2ccc(cc2)O)/C1=O
InChi: 1S/C20H16N2O4S/c1-2-10-22-18(24)17(11-13-6-8-16(23)9-7-13)27-20(22)21-15-5-3-4-14(12-15)19(25)26/h2-9,11-12,23H,1,10H2,(H,25,26)/b17-11-,21-20-
InChiKey: IBMPRVWUMIDHFQ-ICSPAVNMSA-N

Network

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Target Layer

Model Organisms

Species	Viewmode
Arabidopsis thaliana
Caenorhabditis elegans
Dictyostelium discoideum
Drosophila melanogaster
Escherichia coli
Homo sapiens
Mus musculus
Oryza sativa
Saccharomyces cerevisiae
Schmidtea mediterranea
Other organisms

TDR Pathogens:

Species	Viewmode
Brugia malayi
Chlamydia trachomatis
Echinococcus granulosus
Entamoeba histolytica
Echinococcus multilocularis
Giardia lamblia
Leishmania major
Loa Loa (eye worm)
Mycobacterium leprae
Mycobacterium tuberculosis
Mycobacterium ulcerans
Onchocerca volvulus
Plasmodium falciparum
Plasmodium vivax
Schistosoma mansoni
Trypanosoma brucei
Trypanosoma cruzi
Toxoplasma gondii
Treponema pallidum
Trichomonas vaginalis
Wolbachia endosymbiont of Brugia malayi

Other Pathogens:

Species	Viewmode
Babesia bovis
Candida albicans
Cryptosporidium hominis
Cryptosporidium parvum
Leishmania braziliensis
Leishmania donovani
Leishmania infantum
Leishmania mexicana
Neospora caninum
Plasmodium berghei
Plasmodium knowlesi
Plasmodium yoelii
Schistosoma japonicum
Trypanosoma brucei gambiense
Trypanosoma congolense
Theileria parva

Compound Layer

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Clustering layers

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Synonyms

No synonyms found for this compound

Targets

Known targets for this compound

Species	Target name	Source	Bibliographic reference
Mycobacterium tuberculosis	Cysteine synthase a CysK1 (O-acetylserine sulfhydrylase A) (O-acetylserine (thiol)-lyase A) (CSASE A)	Starlite/ChEMBL	References

Predicted pathogen targets for this compound

By orthology

Species	Potential target	Known druggable target/s	Ortholog Group
Mycobacterium tuberculosis	Cysteine synthase a CysK1 (O-acetylserine sulfhydrylase A) (O-acetylserine (thiol)-lyase A) (CSASE A)	Get druggable targets OG5_126662	All targets in OG5_126662
Candida albicans	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Toxoplasma gondii	pyridoxal-phosphate dependent superfamily protein	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Brugia malayi	Pyridoxal-phosphate dependent enzyme family protein	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania braziliensis	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Entamoeba histolytica	cysteine synthase A, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Loa Loa (eye worm)	cbs-prov protein	Get druggable targets OG5_126662	All targets in OG5_126662
Candida albicans	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania mexicana	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium leprae	Probable cystathionine beta-synthase CBS (Serine sulfhydrase) (Beta-thionase) (Hemoprotein H-450)	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania donovani	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	serine sulfhydrylase	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	threonine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania donovani	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania major	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania braziliensis	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium ulcerans	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Schistosoma mansoni	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania mexicana	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	serine sulfhydrylase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	serine sulfhydrylase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma brucei gambiense	cystathionine beta-synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania infantum	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium ulcerans	cysteine synthase a CysK1	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	serine sulfhydrylase	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania infantum	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Schistosoma japonicum	expressed protein	Get druggable targets OG5_126662	All targets in OG5_126662
Entamoeba histolytica	cysteine synthase A, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium leprae	PROBABLE CYSTEINE SYNTHASE A CYSK (O-ACETYLSERINE SULFHYDRYLASE A) (O-ACETYLSERINE (THIOL)-LYASE A) (CSASE A)	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma brucei	cystathionine beta-synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium tuberculosis	Cysteine synthase B CysM (CSASE B) (O-phosphoserine sulfhydrylase B) (O-phosphoserine (thiol)-lyase B)	Get druggable targets OG5_126662	All targets in OG5_126662
Toxoplasma gondii	CBS family protein	Get druggable targets OG5_126662	All targets in OG5_126662
Leishmania major	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium ulcerans	cysteine synthase B	Get druggable targets OG5_126662	All targets in OG5_126662
Mycobacterium tuberculosis	Probable cystathionine beta-synthase Cbs (serine sulfhydrase) (beta-thionase) (hemoprotein H-450)	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	threonine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Entamoeba histolytica	cysteine synthase A, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Trichomonas vaginalis	cysteine synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662
Brugia malayi	cbs-prov protein	Get druggable targets OG5_126662	All targets in OG5_126662
Neospora caninum	Protein ZC373.1, partially confirmed by transcript evidence, related	Get druggable targets OG5_126662	All targets in OG5_126662
Schistosoma mansoni	cysteine synthase	Get druggable targets OG5_126662	All targets in OG5_126662
Trypanosoma cruzi	cystathionine beta-synthase, putative	Get druggable targets OG5_126662	All targets in OG5_126662

By sequence similarity to non orthologous known druggable targets

Species	Potential target	Known druggable target	Length	Alignment span	Identity
Brugia malayi	Pyridoxal-phosphate dependent enzyme family protein	Cysteine synthase a CysK1 (O-acetylserine sulfhydrylase A) (O-acetylserine (thiol)-lyase A) (CSASE A)	310 aa	337 aa	24.0 %

Ranking Plot

Putative Targets List

Species	Potential target	Raw	Global	Species
Echinococcus granulosus	tm gpcr rhodopsin	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	pyroglutamylated rfamide peptide receptor	0.0011	0	0.5
Onchocerca volvulus	Neuropeptide F receptor homolog	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus granulosus	g protein linked acetylcholine receptor gar 2a	0.0011	0	0.5
Trypanosoma cruzi	cystathionine beta-synthase	0.0012	1	0.5
Trichomonas vaginalis	threonine synthase, putative	0.0012	1	0.5
Onchocerca volvulus		0.0011	0	0.5
Toxoplasma gondii	pyridoxal-phosphate dependent superfamily protein	0.0012	1	0.5
Echinococcus multilocularis	fmrfamide receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	neuropeptide Y receptor	0.0011	0	0.5
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5
Echinococcus granulosus	g protein coupled receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus granulosus	fmrfamide receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus granulosus	G protein coupled receptor 139	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Trypanosoma cruzi	cystathionine beta-synthase	0.0012	1	0.5
Echinococcus multilocularis	g protein coupled receptor	0.0011	0	0.5
Echinococcus granulosus	neuropeptides capa receptor	0.0011	0	0.5
Trypanosoma cruzi	serine sulfhydrylase	0.0012	1	0.5
Echinococcus granulosus	7tm_1 domain containing protein	0.0011	0	0.5
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5
Echinococcus multilocularis	G-protein coupled receptor, putative	0.0011	0	0.5
Echinococcus granulosus	g protein linked acetylcholine receptor gar 2a	0.0011	0	0.5
Echinococcus granulosus	neuropeptide s receptor	0.0011	0	0.5
Loa Loa (eye worm)	cbs-prov protein	0.0012	1	1
Echinococcus multilocularis	peptide (allatostatin:somatostatin)	0.0011	0	0.5
Echinococcus multilocularis	g-protein coupled receptor	0.0011	0	0.5
Echinococcus multilocularis	tm gpcr rhodopsin gpcr rhodopsin superfamily	0.0011	0	0.5
Echinococcus multilocularis	serotonin receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	tachykinin peptides receptor 99D	0.0011	0	0.5
Mycobacterium tuberculosis	Probable cystathionine beta-synthase Cbs (serine sulfhydrase) (beta-thionase) (hemoprotein H-450)	0.0012	1	0.5
Schistosoma mansoni	cysteine synthase	0.0012	1	1
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5
Echinococcus granulosus	pyroglutamylated rfamide peptide receptor	0.0011	0	0.5
Mycobacterium leprae	Probable cystathionine beta-synthase CBS (Serine sulfhydrase) (Beta-thionase) (Hemoprotein H-450)	0.0012	1	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	dro:myosuppressin receptor	0.0011	0	0.5
Echinococcus granulosus	growth hormone secretagogue receptor type 1	0.0011	0	0.5
Mycobacterium ulcerans	cysteine synthase a CysK1	0.0012	1	0.5
Echinococcus granulosus	tachykinin peptides receptor 99D	0.0011	0	0.5
Echinococcus granulosus	neuropeptide receptor	0.0011	0	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Entamoeba histolytica	cysteine synthase A, putative	0.0012	1	0.5
Echinococcus multilocularis	neuropeptides capa receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	fmrfamide receptor	0.0011	0	0.5
Echinococcus granulosus	alpha 1A adrenergic receptor	0.0011	0	0.5
Echinococcus granulosus	g protein coupled receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Mycobacterium tuberculosis	Cysteine synthase B CysM (CSASE B) (O-phosphoserine sulfhydrylase B) (O-phosphoserine (thiol)-lyase B)	0.0012	1	0.5
Entamoeba histolytica	cysteine synthase A, putative	0.0012	1	0.5
Echinococcus multilocularis	tm gpcr rhodopsin gpcr rhodopsin superfamily	0.0011	0	0.5
Echinococcus granulosus	dro:myosuppressin receptor	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide FF receptor 2	0.0011	0	0.5
Echinococcus granulosus	tm gpcr rhodopsin	0.0011	0	0.5
Mycobacterium leprae	PROBABLE CYSTEINE SYNTHASE A CYSK (O-ACETYLSERINE SULFHYDRYLASE A) (O-ACETYLSERINE (THIOL)-LYASE A) (CSASE A)	0.0012	1	0.5
Echinococcus multilocularis	g protein linked acetylcholine receptor gar 2a	0.0011	0	0.5
Echinococcus granulosus	tm gpcr rhodopsin	0.0011	0	0.5
Echinococcus granulosus	neuropeptides capa receptor	0.0011	0	0.5
Echinococcus granulosus	neuropeptide receptor A26	0.0011	0	0.5
Echinococcus multilocularis	g protein coupled receptor	0.0011	0	0.5
Leishmania major	cysteine synthase	0.0012	1	0.5
Trypanosoma cruzi	cysteine synthase, putative	0.0012	1	0.5
Echinococcus multilocularis	tm gpcr rhodopsin gpcr rhodopsin superfamily	0.0011	0	0.5
Echinococcus multilocularis	G protein coupled receptor 139	0.0011	0	0.5
Mycobacterium ulcerans	cystathionine beta-synthase	0.0012	1	0.5
Echinococcus granulosus	neuropeptide Y receptor	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus granulosus	thyrotropin releasing hormone receptor	0.0011	0	0.5
Mycobacterium tuberculosis	Cysteine synthase a CysK1 (O-acetylserine sulfhydrylase A) (O-acetylserine (thiol)-lyase A) (CSASE A)	0.0012	1	0.5
Echinococcus granulosus	neuropeptide Y receptor	0.0011	0	0.5
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	thyrotropin releasing hormone receptor	0.0011	0	0.5
Echinococcus multilocularis	growth hormone secretagogue receptor type 1	0.0011	0	0.5
Onchocerca volvulus	Dopamine\/Ecdysteroid receptor homolog	0.0011	0	0.5
Echinococcus multilocularis	alpha 1A adrenergic receptor	0.0011	0	0.5
Schistosoma mansoni	cysteine synthase	0.0012	1	1
Onchocerca volvulus		0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Entamoeba histolytica	cysteine synthase A, putative	0.0012	1	0.5
Trypanosoma brucei	cystathionine beta-synthase, putative	0.0012	1	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Trypanosoma cruzi	cystathionine beta-synthase	0.0012	1	0.5
Onchocerca volvulus		0.0011	0	0.5
Trypanosoma cruzi	cysteine synthase, putative	0.0012	1	0.5
Echinococcus granulosus	neuropeptide FF receptor 2	0.0011	0	0.5
Echinococcus granulosus	hypothetical protein	0.0011	0	0.5
Echinococcus granulosus	sex peptide receptor	0.0011	0	0.5
Echinococcus multilocularis	somatostatin receptor	0.0011	0	0.5
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5
Trypanosoma cruzi	serine sulfhydrylase	0.0012	1	0.5
Echinococcus granulosus	orexin receptor type 2	0.0011	0	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	orexin receptor type 2	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	g protein linked acetylcholine receptor gar 2a	0.0011	0	0.5
Echinococcus multilocularis	7TM GPCR, rhodopsin	0.0011	0	0.5
Echinococcus granulosus	allatostatin A receptor	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide Y receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	biogenic amine (5HT) receptor	0.0011	0	0.5
Echinococcus granulosus	serotonin receptor	0.0011	0	0.5
Leishmania major	cystathionine beta-synthase	0.0012	1	0.5
Echinococcus granulosus	G protein coupled 5 hydroxytryptamine receptor	0.0011	0	0.5
Echinococcus granulosus	g protein coupled receptor	0.0011	0	0.5
Echinococcus multilocularis	serotonin receptor	0.0011	0	0.5
Echinococcus granulosus	g-protein coupled receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide receptor A26	0.0011	0	0.5
Echinococcus granulosus	somatostatin receptor	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide receptor	0.0011	0	0.5
Trypanosoma cruzi	serine sulfhydrylase	0.0012	1	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	neuropeptide s receptor	0.0011	0	0.5
Echinococcus granulosus	neuropeptide receptor	0.0011	0	0.5
Mycobacterium ulcerans	cysteine synthase B	0.0012	1	0.5
Echinococcus multilocularis	sex peptide receptor	0.0011	0	0.5
Echinococcus multilocularis	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus multilocularis	allatostatin A receptor	0.0011	0	0.5
Toxoplasma gondii	CBS family protein	0.0012	1	0.5
Brugia malayi	cbs-prov protein	0.0012	1	1
Echinococcus multilocularis	G protein coupled 5 hydroxytryptamine receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus granulosus	biogenic amine 5HT receptor	0.0011	0	0.5
Trypanosoma cruzi	cystathionine beta-synthase, putative	0.0012	1	0.5
Trypanosoma cruzi	cystathionine beta-synthase, putative	0.0012	1	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus granulosus	biogenic amine 5HT receptor	0.0011	0	0.5
Trypanosoma cruzi	serine sulfhydrylase	0.0012	1	0.5
Echinococcus multilocularis	g protein coupled receptor	0.0011	0	0.5
Onchocerca volvulus		0.0011	0	0.5
Echinococcus multilocularis	neuropeptides capa receptor	0.0011	0	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus granulosus	peptide allatostatin:somatostatin	0.0011	0	0.5
Trichomonas vaginalis	threonine synthase, putative	0.0012	1	0.5
Trypanosoma cruzi	cystathionine beta-synthase, putative	0.0012	1	0.5
Echinococcus granulosus	rhodopsin orphan GPCR	0.0011	0	0.5
Echinococcus granulosus	7TM GPCR rhodopsin	0.0011	0	0.5
Trichomonas vaginalis	cysteine synthase, putative	0.0012	1	0.5

Activities

Activity type	Activity value	Assay description	Source	Reference
IC50 (binding)	= 110.4 nM	Inhibition of Mycobacterium tuberculosis N-terminal His-6-tagged recombinant CysK1 expressed in Escherichia coli BL21 (DE3) after 20 mins by spectrophotometric analysis	ChEMBL.	23879381
Inhibition (binding)	= 92.3 %	Inhibition of Mycobacterium tuberculosis N-terminal His-6-tagged recombinant CysK1 expressed in Escherichia coli BL21 (DE3) at 500 nM after 20 mins by spectrophotometric analysis	ChEMBL.	23879381

Phenotypes

Whole-cell/tissue/organism interactions

We have no records of whole-cell/tissue assays done with this compound What does this mean?

Many chemical entities in TDR Targets come from high-throughput screenings with whole cells or tissue samples, and not all assayed compounds have been tested against a single a single target protein, probably because they get ruled out during screening process. Even if these compounds may have not been of interest in the original screening, they may come as interesting leads for other screening assays. Furthermore, we may be able to propose drug-target associations using chemical similarities and network patterns.

Annotated phenotypes:

We have no manually annotated phenotypes for this drug. What does this mean? / Care to help?

In TDR Targets, information about phenotypes that are caused by drugs, or by genetic manipulation of cells (e.g. gene knockouts or knockdowns) is manually curated from the literature. These descriptions help to describe the potential of the target for drug development. If no information is available for this gene or if the information is incomplete, this may mean that i) the papers containing this information either appeared after the curation effort for this organism was carried out or they were inadvertently missed by curators; or that ii) the curation effort for this organism has not yet started.

In any case, if you have information about papers containing relevant validation data for this target, please log in using your TDR Targets username and password and send them to us using the corresponding form in this page (only visible to registered users) or contact us.

External resources for this compound

ChEMBL: CHEMBL2418548

Bibliographic References

1 literature reference was collected for this gene.

If you have references for this compound, please enter them in a user comment (below) or Contact us.

Detailed information for compound 1767827