Detailed information for compound 957120

Basic information

Technical information
  • TDR Targets ID: 957120
  • Name: 2-[(6-chloro-1H-imidazo[4,5-b]pyridin-2-yl)su lfanyl]-N-(3,4-dimethoxyphenyl)acetamide
  • MW: 378.833 | Formula: C16H15ClN4O3S
  • H donors: 2 H acceptors: 3 LogP: 3.07 Rotable bonds: 7
    Rule of 5 violations (Lipinski): 1
  • SMILES: COc1cc(ccc1OC)NC(=O)CSc1nc2c([nH]1)ncc(c2)Cl
  • InChi: 1S/C16H15ClN4O3S/c1-23-12-4-3-10(6-13(12)24-2)19-14(22)8-25-16-20-11-5-9(17)7-18-15(11)21-16/h3-7H,8H2,1-2H3,(H,19,22)(H,18,20,21)
  • InChiKey: JPROKJDNDHXYFM-UHFFFAOYSA-N  


Substructure search Similarity search

Network

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Synonyms

  • 2-[(6-chloro-1H-imidazo[4,5-b]pyridin-2-yl)thio]-N-(3,4-dimethoxyphenyl)acetamide
  • 2-[(6-chloro-1H-imidazo[4,5-b]pyridin-2-yl)sulfanyl]-N-(3,4-dimethoxyphenyl)ethanamide
  • ZINC05310416

Targets

Known targets for this compound

Species Target name Source Bibliographic reference
Homo sapiens ectonucleotide pyrophosphatase/phosphodiesterase 1 Starlite/ChEMBL References

Predicted pathogen targets for this compound

By orthology
Species Potential target Known druggable target/s Ortholog Group
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma japonicum Ectonucleotide pyrophosphatase/phosphodiesterase family member 5 precursor, putative Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Loa Loa (eye worm) hypothetical protein Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma japonicum Ectonucleotide pyrophosphatase/phosphodiesterase family member 5 precursor, putative Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Onchocerca volvulus Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Loa Loa (eye worm) hypothetical protein Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Loa Loa (eye worm) hypothetical protein Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Brugia malayi Type I phosphodiesterase / nucleotide pyrophosphatase family protein Get druggable targets OG5_127260 All targets in OG5_127260
Schistosoma japonicum ko:K01175 ENPP5, LOC481824; ectonucleotide pyrophosphatase/phosphodiesterase 5 (putative function) [EC:3.1.-.-], putative Get druggable targets OG5_127260 All targets in OG5_127260
Onchocerca volvulus Get druggable targets OG5_127260 All targets in OG5_127260
Candida albicans similar to human plasma membrane glycoprotein PCI Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus granulosus ectonucleotide Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
Candida albicans similar to human plasma membrane glycoprotein PCI Get druggable targets OG5_127260 All targets in OG5_127260
Loa Loa (eye worm) hypothetical protein Get druggable targets OG5_127260 All targets in OG5_127260
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase Get druggable targets OG5_127260 All targets in OG5_127260
By sequence similarity to non orthologous known druggable targets
No druggable targets predicted by sequence similarity
Obtained from network model

Ranking Plot

Putative Targets List

Species Potential target Raw Global Species
Schistosoma mansoni hypothetical protein 0.0041 0.0272 0.0302
Schistosoma mansoni thyroid hormone receptor 0.0023 0.0073 0.0067
Schistosoma mansoni FTZ-F1 nuclear receptor-like protein 0.0023 0.0073 0.0067
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.03
Brugia malayi nuclear receptor NHR-88 0.0023 0.0073 0.0238
Schistosoma mansoni histone-lysine n-methyltransferase setb1 0.0034 0.0193 0.0209
Echinococcus granulosus nuclear receptor 2DBD gamma 0.0023 0.0073 0.0067
Echinococcus multilocularis FTZ F1 alpha 0.0023 0.0073 0.0057
Brugia malayi steroid hormone receptor 0.0023 0.0073 0.0238
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.03
Echinococcus multilocularis FTZ F1 nuclear receptor protein 0.0023 0.0073 0.0057
Echinococcus multilocularis thyroid hormone receptor alpha 0.0023 0.0073 0.0057
Loa Loa (eye worm) hypothetical protein 0.0034 0.0199 0.0652
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Schistosoma mansoni histone-lysine n-methyltransferase suv9 0.0034 0.0193 0.0209
Brugia malayi Intermediate filament tail domain containing protein 0.003 0.0158 0.0518
Toxoplasma gondii LsmAD domain-containing protein 0.0028 0.0136 0.449
Schistosoma mansoni histone-lysine n-methyltransferase setb1 0.0034 0.0193 0.0209
Brugia malayi Corticotropin releasing factor receptor 2 precursor, putative 0.0056 0.0444 0.1454
Echinococcus granulosus intermediate filament protein 0.003 0.0158 0.0168
Brugia malayi Nuclear hormone receptor family member nhr-19 0.0023 0.0073 0.0238
Onchocerca volvulus 0.004 0.027 0.0725
Plasmodium falciparum ataxin-2 like protein, putative 0.0028 0.0136 0.5
Onchocerca volvulus 0.003 0.0158 0.0313
Loa Loa (eye worm) latrophilin receptor protein 2 0.0018 0.0016 0.0052
Loa Loa (eye worm) nuclear hormone receptor family member nhr-41 0.0023 0.0073 0.0238
Loa Loa (eye worm) hypothetical protein 0.0038 0.0246 0.0805
Echinococcus granulosus Endonuclease G 0.0024 0.0089 0.0087
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Entamoeba histolytica hypothetical protein 0.0041 0.0272 0.5
Loa Loa (eye worm) nuclear hormone receptor family member nhr-40 0.0023 0.0073 0.0238
Leishmania major hypothetical protein, conserved 0.0028 0.0136 1
Schistosoma mansoni Guanine nucleotide-binding protein G(s) subunit alpha (Adenylate cyclase-stimulating G alpha protein) 0.0051 0.0392 0.0444
Loa Loa (eye worm) intermediate filament protein 0.003 0.0158 0.0518
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase 0.004 0.027 0.03
Brugia malayi ecdysteroid receptor 0.0023 0.0073 0.0238
Brugia malayi latrophilin 2 splice variant baaae 0.0038 0.0246 0.0805
Schistosoma mansoni lamin 0.003 0.0158 0.0168
Echinococcus granulosus Nuclear hormone receptor family member nhr 41 0.0023 0.0073 0.0067
Loa Loa (eye worm) pre-SET domain-containing protein family protein 0.0234 0.2431 0.7969
Schistosoma mansoni photoreceptor-specific nuclear receptor related 0.0023 0.0073 0.0067
Brugia malayi nuclear hormone receptor 0.0023 0.0073 0.0238
Loa Loa (eye worm) hypothetical protein 0.0034 0.0193 0.0631
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase 0.004 0.027 0.03
Echinococcus granulosus COUP TF:Svp nuclear hormone receptor 0.0023 0.0073 0.0067
Echinococcus granulosus 5'partial|histone lysine N methyltransferase SETDB2 0.0032 0.0179 0.0192
Loa Loa (eye worm) hypothetical protein 0.003 0.0152 0.0497
Onchocerca volvulus 0.0266 0.2792 1
Schistosoma mansoni transcription factor LCR-F1 0.0041 0.0272 0.0302
Echinococcus granulosus nuclear receptor 2DBD gamma 0.0023 0.0073 0.0067
Brugia malayi Latrophilin receptor protein 2 0.0018 0.0016 0.0052
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Echinococcus granulosus lamin 0.003 0.0158 0.0168
Loa Loa (eye worm) nuclear Hormone Receptor family member 0.0023 0.0073 0.0238
Brugia malayi Nuclear hormone receptor family member nhr-25 0.0023 0.0073 0.0238
Echinococcus multilocularis lamin dm0 0.003 0.0158 0.0142
Brugia malayi hypothetical protein 0.0041 0.0272 0.089
Brugia malayi Nuclear hormone receptor family member nhr-3 0.0023 0.0073 0.0238
Onchocerca volvulus 0.004 0.027 0.0725
Brugia malayi Nuclear hormone receptor family member nhr-25 0.0023 0.0073 0.0238
Loa Loa (eye worm) nuclear hormone receptor family member nhr-31 0.0023 0.0073 0.0238
Echinococcus multilocularis Endonuclease G 0.0024 0.0089 0.0074
Brugia malayi Nuclear hormone receptor-like 1 0.0023 0.0073 0.0238
Schistosoma mansoni microtubule-associated protein tau 0.0775 0.8483 1
Onchocerca volvulus 0.0087 0.0794 0.2652
Echinococcus multilocularis histone lysine methyltransferase setb histone lysine methyltransferase eggless 0.0034 0.0193 0.0177
Echinococcus multilocularis Basic leucine zipper (bZIP) transcription 0.0041 0.0272 0.0256
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.03
Schistosoma mansoni intermediate filament proteins 0.003 0.0158 0.0168
Brugia malayi Steroid receptor seven-up type 2 0.0023 0.0073 0.0238
Entamoeba histolytica hypothetical protein 0.0041 0.0272 0.5
Plasmodium falciparum ataxin-2 like protein, putative 0.0028 0.0136 0.5
Brugia malayi Ligand-binding domain of nuclear hormone receptor family protein 0.0023 0.0073 0.0238
Echinococcus multilocularis ecdysone induced protein 78C 0.0023 0.0073 0.0057
Brugia malayi Nuclear hormone receptor family member nhr-40 0.0023 0.0073 0.0238
Loa Loa (eye worm) hypothetical protein 0.003 0.0158 0.0518
Echinococcus granulosus FTZ F1 alpha 0.0023 0.0073 0.0067
Echinococcus multilocularis nuclear receptor 2DBD gamma 0.0023 0.0073 0.0057
Schistosoma mansoni hypothetical protein 0.0087 0.0794 0.0919
Brugia malayi calcium-independent alpha-latrotoxin receptor 2, putative 0.0018 0.0016 0.0052
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Brugia malayi Nuclear hormone receptor family member nhr-14 0.0023 0.0073 0.0238
Echinococcus multilocularis histone lysine N methyltransferase SETMAR 0.0034 0.0193 0.0177
Trypanosoma brucei PAB1-binding protein , putative 0.0028 0.0136 1
Echinococcus granulosus Basic leucine zipper bZIP transcription 0.0041 0.0272 0.0302
Loa Loa (eye worm) nuclear hormone receptor family member nhr-14 0.0023 0.0073 0.0238
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Trichomonas vaginalis set domain proteins, putative 0.0266 0.2792 0.5
Brugia malayi hypothetical protein 0.0028 0.0136 0.0445
Schistosoma mansoni nuclear receptor 2DBD-gamma 0.0023 0.0073 0.0067
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Echinococcus multilocularis microtubule associated protein 2 0.0775 0.8483 0.848
Loa Loa (eye worm) GTP-binding regulatory protein Gs alpha-S chain 0.0051 0.0392 0.1285
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Loa Loa (eye worm) steroid hormone receptor 0.0023 0.0073 0.0238
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.03
Schistosoma mansoni thyroid hormone receptor 0.0023 0.0073 0.0067
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Brugia malayi Nuclear hormone receptor family member nhr-31 0.0023 0.0073 0.0238
Schistosoma mansoni nuclear hormone receptor nor-1/nor-2 0.0023 0.0073 0.0067
Echinococcus granulosus histone lysine methyltransferase setb 0.0034 0.0193 0.0209
Onchocerca volvulus 0.003 0.0158 0.0313
Echinococcus multilocularis nuclear receptor 2DBD gamma 0.0023 0.0073 0.0057
Echinococcus granulosus guanine nucleotide binding protein Gs subunit 0.0051 0.0392 0.0444
Brugia malayi intermediate filament protein 0.003 0.0158 0.0518
Schistosoma mansoni Guanine nucleotide-binding protein G(s) subunit alpha (Adenylate cyclase-stimulating G alpha protein) 0.0051 0.0392 0.0444
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Schistosoma mansoni lamin 0.003 0.0158 0.0168
Loa Loa (eye worm) intermediate filament tail domain-containing protein 0.003 0.0158 0.0518
Brugia malayi Nuclear hormone receptor family member nhr-1 0.0023 0.0073 0.0238
Toxoplasma gondii histone lysine methyltransferase SET/SUV39 0.0034 0.0193 1
Schistosoma mansoni histone-lysine n-methyltransferase setb1 0.0034 0.0193 0.0209
Brugia malayi Pre-SET motif family protein 0.0234 0.2431 0.7969
Echinococcus granulosus FTZ F1 nuclear receptor protein 0.0023 0.0073 0.0067
Plasmodium vivax SET domain protein, putative 0.0034 0.0193 1
Schistosoma mansoni nuclear hormone receptor 0.0023 0.0073 0.0067
Schistosoma mansoni steroid hormone receptor ad4bp 0.0023 0.0073 0.0067
Loa Loa (eye worm) hypothetical protein 0.0056 0.0444 0.1454
Brugia malayi Thrombospondin type 1 domain containing protein 0.0087 0.0794 0.2602
Echinococcus multilocularis guanine nucleotide binding protein G(s) subunit 0.0051 0.0392 0.0377
Echinococcus granulosus ecdysone induced protein 78C 0.0023 0.0073 0.0067
Echinococcus granulosus microtubule associated protein 2 0.0775 0.8483 1
Echinococcus granulosus guanine nucleotide binding protein Gs subunit 0.0051 0.0392 0.0444
Brugia malayi GTP-binding regulatory protein Gs alpha-S chain, putative 0.0051 0.0392 0.1285
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Brugia malayi Ligand-binding domain of nuclear hormone receptor family protein 0.0023 0.0073 0.0238
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.0034 0.0199 0.0216
Echinococcus multilocularis Nuclear hormone receptor family member nhr 41 0.0023 0.0073 0.0057
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.0255
Echinococcus multilocularis musashi 0.003 0.0158 0.0142
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase 0.004 0.027 0.03
Echinococcus granulosus ectonucleotide pyrophosphatase:phosphodiesterase 0.004 0.027 0.03
Brugia malayi Calcitonin receptor-like protein seb-1 0.0056 0.0444 0.1454
Brugia malayi Nuclear hormone receptor family member nhr-19 0.0023 0.0073 0.0238
Loa Loa (eye worm) nuclear hormone receptor family member nhr-49 0.0023 0.0073 0.0238
Brugia malayi Nuclear hormone receptor family member nhr-49 0.0023 0.0073 0.0238
Trypanosoma cruzi PAB1-binding protein , putative 0.0028 0.0136 1
Loa Loa (eye worm) hypothetical protein 0.004 0.027 0.0885
Echinococcus multilocularis ectonucleotide pyrophosphatase:phosphodiesterase 0.0034 0.0199 0.0183
Loa Loa (eye worm) hypothetical protein 0.0023 0.0073 0.0238
Loa Loa (eye worm) nuclear hormone receptor family member nhr-1 0.0023 0.0073 0.0238
Echinococcus multilocularis ectonucleotide 0.0034 0.0199 0.0183
Echinococcus granulosus ectonucleotide 0.0034 0.0199 0.0216
Loa Loa (eye worm) hypothetical protein 0.0028 0.0136 0.0445
Schistosoma mansoni endonuclease related 0.0024 0.0089 0.0087
Echinococcus granulosus lamin dm0 0.003 0.0158 0.0168
Brugia malayi Nuclear hormone receptor family member nhr-41 0.0023 0.0073 0.0238
Brugia malayi hypothetical protein 0.0018 0.0023 0.0076
Entamoeba histolytica hypothetical protein 0.0041 0.0272 0.5
Schistosoma mansoni retinoic acid receptor RXR 0.0023 0.0073 0.0067
Echinococcus multilocularis guanine nucleotide binding protein G(s) subunit 0.0051 0.0392 0.0377
Echinococcus granulosus retinoic acid receptor rxr beta a 0.0023 0.0073 0.0067
Schistosoma mansoni RAR-like nuclear receptor 0.0023 0.0073 0.0067
Loa Loa (eye worm) thrombospondin type 1 domain-containing protein 0.0087 0.0794 0.2602
Schistosoma mansoni hypothetical protein 0.0038 0.0246 0.0271
Brugia malayi endonuclease G, mitochondrial precursor 0.0024 0.0089 0.0293
Loa Loa (eye worm) pigment dispersing factor receptor c 0.0056 0.0444 0.1454
Loa Loa (eye worm) hypothetical protein 0.004 0.027 0.0885
Trypanosoma cruzi PAB1-binding protein , putative 0.0028 0.0136 1
Schistosoma mansoni Tr4/Tr2 (homologue) 0.0023 0.0073 0.0067
Brugia malayi Type I phosphodiesterase / nucleotide pyrophosphatase family protein 0.004 0.027 0.0885
Loa Loa (eye worm) hypothetical protein 0.0018 0.0016 0.0052
Brugia malayi Pre-SET motif family protein 0.0034 0.0193 0.0631
Schistosoma mansoni Guanine nucleotide-binding protein G(s) subunit alpha (Adenylate cyclase-stimulating G alpha protein) 0.0051 0.0392 0.0444
Schistosoma mansoni ectonucleotide pyrophosphatase/phosphodiesterase 0.004 0.027 0.03
Brugia malayi photoreceptor-specific nuclear receptor 0.0289 0.3051 1
Echinococcus granulosus hepatocyte nuclear factor 4 alpha 0.0023 0.0073 0.0067
Schistosoma mansoni coup transcription factor 0.0023 0.0073 0.0067
Entamoeba histolytica hypothetical protein 0.0041 0.0272 0.5
Loa Loa (eye worm) hypothetical protein 0.0289 0.3051 1
Onchocerca volvulus 0.0034 0.0193 0.0441
Echinococcus multilocularis hepatocyte nuclear factor 4 alpha 0.0023 0.0073 0.0057
Echinococcus multilocularis lamin 0.003 0.0158 0.0142
Loa Loa (eye worm) hypothetical protein 0.004 0.027 0.0885
Echinococcus multilocularis COUP TF:Svp nuclear hormone receptor 0.0023 0.0073 0.0057
Schistosoma mansoni retinoid-x-receptor (RXR) 0.0023 0.0073 0.0067

Activities

Activity type Activity value Assay description Source Reference
Ki (binding) = 0.137 uM Competitive inhibition of human NPP1 using p-Nph-5'-TMP as substrate by Lineweaver-Burk plot analysis ChEMBL. 25372276
Ki (binding) = 0.137 uM Inhibition of human NPP1 using p-Nph-5'-TMP as substrate ChEMBL. 25372276
Ki (binding) = 9.78 uM Competitive inhibition of human NPP1 using ATP as substrate by Lineweaver-Burk plot analysis ChEMBL. 25372276
Ratio Ki (binding) = 100 Ratio of Ki for human NPP1 using ATP as substrate to Ki for human NPP1 using p-Nph-5'-TMP as substrate ChEMBL. 25372276

Phenotypes

Whole-cell/tissue/organism interactions

We have no records of whole-cell/tissue assays done with this compound What does this mean?

Many chemical entities in TDR Targets come from high-throughput screenings with whole cells or tissue samples, and not all assayed compounds have been tested against a single a single target protein, probably because they get ruled out during screening process. Even if these compounds may have not been of interest in the original screening, they may come as interesting leads for other screening assays. Furthermore, we may be able to propose drug-target associations using chemical similarities and network patterns.

Annotated phenotypes:

We have no manually annotated phenotypes for this drug. What does this mean? / Care to help?
In TDR Targets, information about phenotypes that are caused by drugs, or by genetic manipulation of cells (e.g. gene knockouts or knockdowns) is manually curated from the literature. These descriptions help to describe the potential of the target for drug development. If no information is available for this gene or if the information is incomplete, this may mean that i) the papers containing this information either appeared after the curation effort for this organism was carried out or they were inadvertently missed by curators; or that ii) the curation effort for this organism has not yet started.
 
In any case, if you have information about papers containing relevant validation data for this target, please log in using your TDR Targets username and password and send them to us using the corresponding form in this page (only visible to registered users) or contact us.

External resources for this compound

Bibliographic References

1 literature reference was collected for this gene.

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